Life habit: lichenized (mutualistic with algal photobionts) thallus: crustose (crustaceous) – episubstratal – unspecified thallus: continuous, diffuse, effuse | granular, granulose, granulate [th] upper surface: grey(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) | grey(ish) white [th] morphol substructures (eg areoles, lobes, branches) upper surface: granulose, granular | subsquamulose ascomata: absent | present ascoma: perithecial, perithecioid – hymenial ascoma [mm]: (low) 0.1 (high) 0.15 (max) 0.3 ascoma: subsessile, subimmersed, adnate, semi-immersed, emergent [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) [ascm, if apoth] margin excipular photobionts: absent [ascm] paraphyses/-oids: absent asci: fissitunicate – verrucarialean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): absent ascospores: (median) 8.0 [asp] length [µm]: (min) 17.0 (low) 21.5 (high) 25.5 (max) 30.0 [asp] width [µm]: (min) 9.0 (low) 12.0 (high) 14.0 (max) 16.5 [asp] septa: present [asp] septa: muriformly septate – muriform, submuriform [asp] pigmentation: hyaline, colourless [asp] perispore, epispore: not apparent secondary metabolites: absent primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts