Life habit: lichenized (mutualistic with algal photobionts) thallus: crustose (crustaceous) – episubstratal – unspecified thallus: granular, granulose, granulate [th] upper surface: black(ish) [th marginal and upper surface] specific structures: absent ascomata: absent | present ascoma: perithecial, perithecioid – hymenial ascoma [mm]: (low) 0.2 (high) 0.3 ascoma: subsessile, subimmersed, adnate, semi-immersed, emergent [ascm] paraphyses/-oids: absent asci: fissitunicate – verrucarialean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): absent ascospores: (median) 8.0 [asp] shape: ellipsoidal [asp] length [µm]: (min) 15.6 (low) 20.0 (high) 30.0 (max) 36.0 [asp] width [µm]: (min) 8.0 (low) 10.0 (high) 15.0 [asp] septa: present [asp] septa: muriformly septate – muriform, submuriform [asp] pigmentation: medium brown, brownish | dark brown | reddish brown [asp] perispore, epispore: not apparent secondary metabolites: absent primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | lichens, lichenized fungi.