Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2002. Lichen Flora of the Greater Sonoran Desert Region. Vol 1.

Thallus: delicately squamulose, forming small aggregates several mm across squamules: 0.1-0.5 mm wide, ± crowded, adnate to ascending, flat or convex, roundish to elongate, more rarely digitiform or divided into coarsely subgranular segments upper surface: grayish brown or green-brown to fawn, green when wet, dull anatomy: paraplectenchymatous throughout, with no distinct cortices; algal cells: distributed throughout most of the thallus lower surface: pale Perithecia: prominent, black, barrel-shaped, up to 0.5 mm broad, surface rough and wrinkled asci: clavate, 2-spored ascospores: strongly muriform, elongate-ellipsoid, hyaline to pale yellow-brown, c. 80-120 x 30-50 micro meters Pycnidia: unknown Spot tests: all negative Secondary metabolites: none detected. Substrate and ecology: over soil, mosses or debris, mainly over calciferous ground, rarely on nutrient-rich bark, mainly upland World distribution: Europe, Macaronesia, North America, Pacific islands Sonoran distribution: very rarely found in Arizona, but certainly overlooked.