Life habit: lichenized (mutualistic with algal photobionts) thallus: foliose (foliaceous), leaf-like thallus: continuous, diffuse, effuse [th] upper surface: grey(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) [th marginal and upper surface] specific structures: present [th margin] cilia, cilioid structures: absent [th upper surface] isidia, isidioid structures: absent [th upper surface] soredia, soralia, soralioid structures: absent [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 20.0 (high) 40.0 (max) 50.0 [th] morphol substructures (eg areoles, squamules): contiguous, coherent (throughout the thallus) [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane [th] lower surface: brown(ish) black | black(ish) brown [th lower surface] specific structures: absent | present [th lower surface] rhizines, rhizoid structures: absent | present ascomata: absent | present ascoma: apothecial, apothecioid – hymenial ascoma: subpedicellate, substipitate, subpedunculate, substalked | pedicellate, stipitate, pedunculate, stalked [ascm, if apoth] disc, mazaedium: concave [ascm, if apoth] disc, mazaedium: red(dish) brown (if pale: orange brown) [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: grey(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) asci: lecanoralean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): present [asc] tholus amyloidity pattern: with amyloid tube, ring (= Collema-, Micarea-, Porpidia-, Psora-types etc) [asp] septa: present [asp] septa: transversely septate [asp] transversal septa: (low) 4.0 (high) 8.0 [asp] pigmentation: hyaline, colourless | pale brown | medium brown, brownish secondary metabolites: present secondary metabolites: phlebic acid A (28-acetoxy-22-hydroxyhopane-23-oic acid) | phlebic acid B (22-hydroxyhopane-23-oic acid) | zeorin (hopane-6α,22-diol) primary photobiont: present secondary photobionts (eg in cephalodia): absent | present primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts.