Ecology: Terricolous or bryophytic. Thallus: Not subdivided parts, areolate (primarily areolate); separate thallus parts thin. Thallus Outline: Indistinct; soon disappearing. Upper Surface: Verrucose or verruculose, lemon (citrine), orange, pale grey, or pale brown, matt, smooth; epruinose; not sorediate. Upper Cortex: Gelatinized. Ascocarps: Soon sessile, strongly constricted at the base. Margin: Lecanorine, verrucose or verruculose, smooth, not flexuose or slightly flexuose, persistent, indistinct or prominent, concolourous with the surrounding thallus; without a thin rim between disk and thalline margin. Disk: Plane or excavate, brownish red, brown, or dark brown, epruinose. Parathecium (proper excipulum): White, dissolving in 10% KOH. Amphithecium (thalline excipulum): Present; crystals small, abundant, dissolving in 10% KOH; photobiont abundant. Amphithecial Cortex: Present, indistinct, eucortical, (35)-40-47-55 µm wide, basally widened, (50)-55-90-120-(130) µm wide at the base; inspersed with oil droplets. Episamma: Without episamma. Epithecium: Apical cells slightly swollen or distinctly swollen, brownish red, insoluble in 10% KOH. Hymenium: Oil inspersed. Hypothecium: White; not inspers or oil inspersed. Ascospores: c. 8 per ascus, ellipsoid, (12)-12.5-14-17.5-(18) µm long, (6)-6.5-8-9.5 µm wide; wall thin, not ornamented. Secondary Metabolites: Atranorin, of the following substance class(es): orcinol depsides and (tri-)terpenoids. Spot Tests: Upper surface: C –, PD –.