Global occurrence: Eurasia – Asia Extratropical | Americas – North America (incl Mexico) | Arctic. Substrate: soil, clay, humus, turf, detritus, dead leaves.  Life habit: lichenized (mutualistic with algal photobionts).  Thallus: crustose (crustaceous) – episubstratal – unspecified; continuous, diffuse, effuse | granular, granulose, granulate; [th] upper surface: yellow(ish) (if dull: buff); [th marginal and upper surface] specific structures: absent; [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 0.2 (high) 0.4; [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane.  Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (low) 0.5 (high) 1.2 (max) 1.8; ascoma: sessile, superficial; [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded | subconvex, slightly convex | convex | distinctly convex | strongly convex, hemispherical, (sub-)globose; [ascm, if apoth] disc, mazaedium: green(ish) | yellow(ish) (if dull: buff) | brown(ish) (if pale: fawn, tan; if mid: cinnamon); [ascm, if apoth] disc, mazaedium: epruinose; [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: yellow(ish) (if dull: buff); [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: hyaline, colourless; [ascm] paraphyses/-oids: present; [ascm] paraphyses/-oid cells width [µm]: (low) 1.5 (high) 3.5; [ascm] paraphyses/-oid apical cells width [µm]: (min) 5.0; [ascm] epihymenium, epithecium: yellow(ish) (if dull: buff) | red(dish) yellow (orange, orange yellow).  Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present

[asc] tholus amyloidity pattern: amyloid with widening axial body towards the apex (= Lecanora-, Parmelia-, Rinodina-types etc); Ascospores: (median) 8.0; [asp] shape: ellipsoidal; [asp] length [µm]: (min) 9.0 (low) 11.0 (high) 15.5 (max) 17.0; [asp] width [µm]: (min) 4.5 (low) 5.0 (high) 6.5 (max) 7.0; [asp] septa: absent – spore lumen unilocular, monolocular | present; [asp] septa: transversely septate; [asp] transversal septa: (median) 1.0; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent.  Secondary metabolites: present, pulvinic acid derivatives.  Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid.  Secondary photobionts (eg in cephalodia): absent.