Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2002. Lichen Flora of the Greater Sonoran Desert Region. Vol 1.

Thallus: continuous, immersed in the substrate, upper surface: whitish to gray, smooth; photobiont: absent Perithecia: hemispherical, black, superficial on the thallus, 0.2-0.3 mm diam. ascomatal wall: black, absent below the hamathecium hamathecium: branched pseudoparaphyses, not anastomosing, dehiscent or becoming unclear; filaments: c. 3 µm wide, not inspersed with oil droplets asci: pyriform, c. 60-70 x 18-22 µm, with 8, irregularly arranged ascospores ascospores: hyaline but overmature ones brownish, clavate, 3-septate (starting with a supramedian euseptum), 25-30 x 6-9 µm; walls: roughly ornamented with warts, without a gelatinous sheath Spot tests: all negative, UV negative Secondary metabolites: none detected. Substrate and ecology: on bark of various woody plants, especially conifers like Abies and Picea World distribution: northern temperate or possibly nearly cosmopolitan Sonoran distribution: recorded in eastern Arizona and Baja California Sur.