Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2002. Lichen Flora of the Greater Sonoran Desert Region. Vol 1.

Life habit: non-lichenized, saprophytic Thallus: absent or immersed in the substrate photobiont: absent Ascomata: perithecial, simple involucrellum: dark brown to black, composed of fungal hyphae interspersed with bark cells ascomatal wall: black, continuous below the hamathecium; true exciple: brown, entire below the perithecial cavity hamathecium: unbranched paraphyses, amyloid (IKI+ blue) asci: bitunicate, cylindrical, ocular chamber rounded, tholus non-amyloid ascospores: brown, ellipsoid, 1-septate with a median distoseptum, 12-17 x 5-9 µm; walls: not ornamented Conidiomata: pycnidial conidia: filiform, hyaline Secondary metabolites: absent Geography: SE U.S.A. to the neotropics Substrate: only on bark. Notes: It is characterized by the brown, 1-septate distoseptate ascospores and the unbranched paraphyses; differing from all other genera of the Pyrenulaceae in the consistently 1-septate, brown ascospores.