Life habit: lichenized (mutualistic with algal photobionts) thallus: crustose (crustaceous) – episubstratal – unspecified [th] upper surface: white(ish) [th marginal and upper surface] specific structures: absent [th] morphol substructures (eg areoles, lobes, branches) upper surface: verrucose, warted ascomata: absent | present ascoma: apothecial, apothecioid – hymenial [ascm, if apoth] disc, mazaedium: epruinose [ascm] paraphyses/-oids: present asci: lecanoralean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): present [asc] tholus amyloidity pattern: amyloid with acuate axial body towards the apex (= Bacidia-, Buellia-, Ramalina-types etc) ascospores: (median) 8.0 [asp] length [µm]: (low) 23.0 (high) 24.0 [asp] width [µm]: (low) 9.0 (high) 13.0 [asp] septa: present [asp] septa: transversely septate [asp] transversal septa: (median) 3.0 [asp] pigmentation: medium brown, brownish secondary metabolites: present secondary metabolites: 6-O-methylarthothelin | capistraton primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts.