Life habit:thallus: crustose (crustaceous) – episubstratal – unspecified | subsquamulose (subsquamulous) thallus: compartimentized – areolate, squamulate [th] upper surface: grey(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) | grey(ish) green [th marginal and upper surface] specific structures: absent | present [th margin] cilia, cilioid structures: absent [th upper surface] isidia, isidioid structures: absent [th upper surface] soredia, soralia, soralioid structures: absent [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 0.08 (high) 0.3 [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane | granulose, granular | verrucose, warted | subsquamulose | cracked, fissured, fractured, rimulose, subrimose ascomata: absent | present ascoma: perithecial, perithecioid – hymenial ascoma [mm]: (low) 0.4 (high) 0.7 (max) 0.8 ascoma: immersed, innate | subsessile, subimmersed, adnate, semi-immersed, emergent [ascm] paraphyses/-oids: absent asci: fissitunicate – verrucarialean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): absent ascospores: (median) 8.0 [asp] shape: ellipsoidal [asp] length [µm]: (min) 39.0 (low) 51.0 (high) 68.0 (max) 81.0 [asp] width [µm]: (min) 22.0 (low) 26.0 (high) 34.0 (max) 40.0 [asp] septa: present [asp] septa: muriformly septate – muriform, submuriform [asp] pigmentation: hyaline, colourless | pale brown | yellowish brown [asp] perispore, epispore: not apparent secondary metabolites: absent primary photobiont: present secondary photobionts (eg in cephalodia): present primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts | rock – siliceous, siliciferous, acidic | rock, stones, pebbles – unspecified.