Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2002. Lichen Flora of the Greater Sonoran Desert Region. Vol 1.

Thallus: foliose, orbicular, rarely irregular, small, less than 2 cm diam., very tightly adnate, often verrucose in center, densely lobate lobes: narrow, 0.3-1.0 (-2.0) mm wide, radiating, usually plane, margins occasionally black, tips sometimes ascending upper surface: gray brown to dark brown, darker at lobe-tips, epruinose or very faintly developed pruina, soredia and isidia absent medulla: white lower surface: dark brown to black centrally, paler to whitish at tips, erhizinate or weakly rhizinate Apothecia: common, up to 2 mm diam., thalline margin entire to crenulate ascospores: brown, ellipsoid, 1-septate, Pachysporaria type, 15-21 x 7-11 µm Pycnidia: ± common, inserted with black weakly protruding tips conidia: filiform, 15-20 x 0.5-1 µm Spot tests: cortex and medulla K- C-, KC-, P- Secondary metabolites: none detected. Substrate and ecology: growing on nutrient rich, riparian trees, such as Prosopis, at low to intermediate elevations World distribution: North America and East Africa Sonoran distribution: occasional in Sonora, Chihuahua, Sinaloa and Baja California Sur. Notes: Its verrucose thallus center, tightly adnate thallus and essential absence of rhizines should separate this species from Phaeophyscia species, that are usually more robust.