Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2004. Lichen Flora of the Greater Sonoran Desert Region. Vol 2.

Thallus: foliose, 2-8 cm in diam., loosely adnate, irregularly lobate lobes: irregular, elongate, plane but inrolled apically, separate, 2-10 mm wide, 125-300 µm thick; apices: rotund, entire, often upturned upper surface: medium gray, usually dull, strongly irregularly wrinkled, not isidiate internal anatomy: with upper and lower cortices consisting of a single layer of irregularly isodiametrical cells 3-10 µm in diam., internally with loosely interwoven chains of Nostoc and hyphae lower surface: light gray, smooth to wrinkled, with scattered tufts of white hairs Apothecia: common, laminal, sessile, 0.5-2 mm wide disc: brown to dark red-brown, concave to plane to convex margin: thalline, concolorous with the thallus or cream-colored, often wrinkled exciple: euparaplectenchymatous, 45-90 µm thick centrally hymenium: hyaline below and thinly yellow to brown above, 90-150 µm tall; paraphyses: unbranched, 2-3 µm wide, slightly inflated apically; subhymenium: pale yellow, 35-70 µm thick asci: cylindrico-clavate, 8-spored ascospores: hyaline, submuriform, 3-5-septate transversely, 0-1-septate longitudinally, ellipsoid to slightly subfusiform, 30-56 x 10-20 µm Pycnidia: not observed Spot tests: all negative Secondary metabolites: none detected. Substrate and ecology: usually on bark of various deciduous trees, rarely on rocks World distribution: especially common in eastern North America, the Adriatic region of Europe, and Australasia Sonoran distribution: Chihuahua and southern part of the Sonoran range in Sinaloa. Note: Leptogium corticola is another example of a species occurring in eastern North America, that apparently follows the Sierra Madre Oriental southwards and the Sierra Madre Occidental northwards into the Sonoran region.