Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2002. Lichen Flora of the Greater Sonoran Desert Region. Vol 1.

lobes: linear, elongate, ± narrow, 0.5-1 (-2) mm wide, subimbricate centrally, ciliate; cilia: prominent, marginal, pale at the base and blackened towards the branched apices upper surface: gray to dark gray without or with a weak pruina, sometimes with laminal cilia; sorediate soredia: in soralia on the underside near the tips, bending the tips upwards and thus appearing labriform (designated capitate by Kurokawa [1962]) upper cortex: prosoplectenchymatous medulla: white above and salmon below lower cortex: lacking lower surface: white but mostly overlaid by a yellow to salmon-colored pigment (K-), arachnoid, rhizinate; rhizines: marginal, simple Apothecia and Pycnidia: not seen Spot tests: cortex K+ yellow, C-, KC-, P+ yellow; medulla K+ yellow, C-, KC-, P- Secondary metabolites: cortex with atranorin and chloroatranorin; medulla with atranorin, zeorin and leucotylin. World distribution: North America from the southern Appalachian Mountains Sonoran distribution: relatively rare in SE Arizona and Baja California Sure, more common in the Sierra Madre Occidental region of Sonora and Chihuahua.