Life habit: lichenized (mutualistic with algal photobionts) thallus: crustose (crustaceous) – episubstratal – unspecified thallus: granular, granulose, granulate [th] upper surface: white(ish) [th upper surface]: epruinose [th marginal and upper surface] specific structures: absent [th] morphol substructures (eg areoles, lobes, branches) upper surface: verrucose, warted [th] medulla amyloidity: absent ascomata: absent | present ascoma: apothecial, apothecioid – hymenial ascoma [mm]: (low) 0.5 (high) 1.0 ascoma: sessile, superficial [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded | subconvex, slightly convex | convex | distinctly convex | strongly convex, hemispherical, (sub-)globose [ascm, if apoth] disc, mazaedium: black(ish) [ascm, if apoth] disc, mazaedium: epruinose [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish) [ascm, if apoth] margin excipular photobionts: absent [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: brown(ish) black [ascm] paraphyses/-oids: present [ascm] paraphyses/-oid cells width [µm]: (low) 1.4 (high) 1.8 [ascm] paraphyses/-oid apical cells width [µm]: (min) 2.5 [ascm] epihymenium, epithecium: brown(ish) black | red(dish) brown (pale: orange brown) asci: lecanoralean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): present [asc] tholus amyloidity pattern: amyloid with acuate axial body towards the apex (= Bacidia-, Buellia-, Ramalina-types etc) ascospores: (median) 8.0 [asp] shape: ellipsoidal [asp] length [µm]: (min) 13.5 (low) 15.0 (high) 21.5 (max) 25.0 [asp] width [µm]: (min) 5.0 (low) 7.0 (high) 10.0 [asp] septa: present [asp] septa: transversely septate [asp] transversal septa: (median) 1.0 [asp] pigmentation: medium brown, brownish | greyish brown [asp] perispore, epispore: not apparent secondary metabolites: present secondary metabolites: atranorin primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts.