Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2002. Lichen Flora of the Greater Sonoran Desert Region. Vol 1.

Life habit: lichenized Thallus: crustose, thin, mostly immersed in substrate upper surface: whitish or pale cortex: lacking photobiont: primary one a trentepohlioid alga, secondary photobiont absent Ascomata: perithecial, almost entirely immersed to sessile, black (rarely pink or whitish), compound, involucrellum: brown-black, hemispherical to globose, surrounding exciple ascomatal wall: black, continuous below the hamathecium exciple: ± globose, colorless or pale brownish hamathecium: branched pseudoparaphyses, anastomosing above the asci, non-amyloid asci: bitunicate, cylindrical, K/I-; apical dome: with a broad ocular chamber surmounted by a hemispherical, meniscus-like structure, non-amyloid, (4-) 8-spored ascospores: uniseriate, hyaline, ellipsoid, 1(-3)-septate with a median euseptum, 11-60 x 5.5-23 µm; walls: ornamented with tiny warts Conidiomata: pycnidial conidia: formed acrogenously, ellipsoid to narrowly ellipsoid, simple, hyaline Secondary metabolites: none detected Geography: mostly temperate on both hemispheres Substrate: mostly on bark, but also on calcareous rock. Notes: It is characterized by its cylindrical asci with an unusual ascal tip structure and ellipsoid, 1-septate ascospores with a warted epispore.