Ecology: Terricolous, bryophytic (especially stumps and logs), or lignicolous; growing sun-exposed, partially shaded, or fully shaded. Thallus: Crustose, squamulose; lobes ascending or horizontal; branches densely aggregated, scattered, loosely aggregated, or clustered; separate thallus parts 3-12 mm long; 1-7 mm wide. Primary Thallus: Persistent. Secondary Thallus: Present, arising from center of the primary thallus, of determinate growth; stipe 15-50 mm high, 1.6-3 mm wide, lime green, homogeneously coloured, corticate, with intact or longitudinally split wall. Thallus Outline: Margin dissected. Upper Surface: Areolated (or broken into irregular "plates"), lime green (maculate); not sorediate. Lower Surface: White or orange (at dying base). Lower Cortex: Absent. Ascocarps: Forming apical at the tip of branches or stipes or along the margin of scyphi, moderately abundant, .5-6 mm in diam.. Disk: Weakly convex to strongly convex; red. Conidiomata: Formed on margins of scyphi. Secondary Metabolites: Squamatic acid and usnic acid, of the following substance class(es): ß-orcinol depsides. Spot Tests: Upper surface: K – (negative), C –, + deep yellow, or + orange, PD –. UV-Fluorescence: Reaction colour: bright bluish white.