|
|
Family: Lecanoraceae
|
MycoBank no. 833381 Diagnosis. Thallus corticolous, verruculose areolate to bullate areolate, faint yellowish green; apothecia sessile, crenate, with olivaceous to blackened, faintly pruinose disc, subhymenium hyaline, hypothecium deep yellowish brown; containing atranorin, thiophanic acid and arthothelin. Type: Ecuador. Galápagos: Santa Cruz, cerca la vía sector Los Gemelos, 0˚38’2.1’’S, 90˚23’37.9’’W, 661 m alt., zona de transición, bosque de Scalesia pedunculata, bosque secundario, perturbado y restaurado, sobre corteza de Psidium galapageium, altura al pecho, 04-Jan-2007, Nugra, F. 279 (CDS 33195–holotype). Description. Thallus corticolous, moderately thickened, verrucose-areolate to bullate-areolate; surface faint yellowish green, smooth, matt, epruinose, lacking soredia; prothallus absent or present as a faint whitish layer extending between the granular areoles, rarely delimiting the thallus outline as a black line. Apothecia numerous, often densely aggregated, circular, 0.4–0.8(-1.1) mm in diam., adnate to sessile, often crowded, distinctly lecanorine, margin smooth to coarsely crenulate, epruinose, concolorous with the thallus, disc plane to slightly convex, deep olivaceous to ±blackened, with a faint, farinose, yellowish green pruina; hymenium hyaline, not inspersed, epihymenium with fine, granular crystals, olive brown, infused by a mixture of fuscous brown (elachista-brown: dissolving in K, HCl± dull greenish, N−) and aeruginose pigments (cinereorufa-green: intensifying in K, HCl+ bluish green, N+ reddish violet), crystals and pigments soluble in K; proper exciple thin, indistinct, lacking crystals; thalline exciple thick, ±corticate, with large crystals only, insoluble in K; subhymenium hyaline, hypothecium deep red-brown (leptocline-brown: K+ orange brown, HCl+ intensely reddish brown, N−); ascospores 8/ascus, simple, ellipsoid, (7.8–)10.3–13.9(–16.7) × (4.9–)5.1–7.4(–9.8) μm (n = 25). Pycnidia not seen. Chemistry. Thallus cortex P± yellowish, K+ yellow, KC+ orange, C+ orange, UV+ pale orange; with atranorin Etymology. Named in honor of Fredy Nugra, who collected the type and only specimen. Ecology and Distribution. At present known only from the type specimen which was collected in upper transition zone forest, on bark of Psidium galapageium. Notes. When clarifying the concept of Vainionora Kalb & Elix (2004) emphasized that species of the genus had a well-developed, pigmented subhymenium, but a colorless hypothecium. In their revision of species of Lecanora with dark pigmented hypothecium Lumbsch et al. (1996) did not distinguish two alternatively pigmented layers (subhymenium and hypothecium) immediately below the hymenium. They considered species with a pigmented hypothecium and filiform conidia to belong to Lecanora s.str. (L. coronulans-group), but species with bacilliform conidia were placed in Vainionora (Lumbsch et al. 1996, p. 286, table 2). |
|