Global occurrence: Eurasia – Asia Extratropical | Eurasia – Europe | Arctic. Substrate: lichens, lichenized fungi | rock – siliceous, siliciferous, acidic | rock, stones, pebbles – unspecified. Life habit: lichenized (mutualistic with algal photobionts) | lichenicolous (in/on lichens). Thallus: crustose (crustaceous) – episubstratal – unspecified; thallus: cracked, fissured, fractured, rimose | compartimentized – areolate, squamulate; [th] upper surface: grey(ish) | white(ish) | brown(ish) grey | | grey(ish) brown | white(ish) brown (beige) | yellow(ish) white (ivory, off-white, cream-coloured); [th upper surface]: epruinose; [th marginal and upper surface] specific structures: present; [th margin] cilia, cilioid structures: absent; [th upper surface] isidia, isidioid structures: absent; [th upper surface] soredia, soralia, soralioid structures: present; [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (min) 0.2 (low) 0.3 (high) 0.5 (max) 1.9; [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane | verrucose, warted | cracked, fissured, fractured, rimulose, subrimose | granulose, granular | uneven, undulate; [th] medulla amyloidity: absent. Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (min) 0.2 (low) 0.4 (high) 1.3 (max) 1.8; ascoma: immersed, innate | subsessile, subimmersed, adnate, semi-immersed, emergent | sessile, superficial; [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded | subconvex, slightly convex | convex; [ascm, if apoth] disc, mazaedium: black(ish) | blue(ish) black; [ascm, if apoth] disc, mazaedium: epruinose; [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish); [ascm, if apoth] margin excipular photobionts: absent; [ascm, if apoth] subhymenial layers, hypothecium, [if perith] basal excipulum: yellow(ish) (if dull: buff) | hyaline, colourless; [ascm] paraphyses/-oids: present; [ascm] paraphyses/-oid cells width [µm]: (median) 1.5; [ascm] paraphyses/-oid apical cells width [µm]: (min) 1.5 (low) 2.0 (high) 4.0 (max) 5.0; [ascm] epihymenium, epithecium: green(ish) | brown(ish) green (olivaceous, olive green). Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: amyloid with widening axial body towards the apex (= Lecanora-, Parmelia-, Rinodina-types etc). Ascospores: (median) 8.0; [asp] shape: ellipsoidal | oblong, oblong-obtuse; [asp] length [µm]: (min) 7.0 (low) 12.0 (high) 15.0; [asp] width [µm]: (min) 4.0 (low) 5.0 (high) 5.5 (max) 6.5; [asp] septa: absent – spore lumen unilocular, monolocular; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent. Conidiomata: absent | present. Conidia: filiform, vermiform; [co] length [µm]: (min) 5.0 (low) 12.0 (high) 29.0 (max) 31.0. Secondary metabolites: present, lobaric acid | miriquidic acid | psoromic acid. Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid. Secondary photobionts (eg in cephalodia): absent.