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General
substrate
calciferous rock
sand
soil
rock (unspecified)
bryophytes
wood
lichens
bark
non-calciferous rock
lifestyle
bryophilous
lichenicolous
lichenized
Ascomata
ascomata*
absent
present
ascomata kind*
perithecioid
apothecioid
Asci
ascus wall thickening
not thickened
thickened
ascus wall amyloidity
present
absent
ascus dehiscence type
prototunicate
unitunicate
bitunicate
lecanoralean
verrucarioid
Ascospores
ascospores septation
absent
present
ascospores wall pigmentation*
pale brownish
medium brownish
hyaline
greyish
dark brownish
Secondary Metabolites [general]
secondary metabolite presence
present
absent
Photobionts
primary photobiont (lichenized/non-lichenized)
present
absent (non-lichenized)
primary photobiont kind
trentepohlioid
cyanobacterioid
chlorococcoid
secondary photobiont - presence/absence
absent (unknown)
present
Grand Canyon National Park
James P. Bennett and Clifford M. Wetmore
Species Count: 252
Muellerella erratica
Acarosporaceae
Acarospora fuscata
Acarospora oligospora
Acarospora privigna
Acarospora schleicheri
Acarospora stapfiana
Acarospora strigata
Glypholecia scabra
Myriospora smaragdula
Pleopsidium chlorophanum
Sarcogyne clavus
Sarcogyne novomexicana
Sarcogyne regularis
Arthoniaceae
Arthonia varians
Caliciaceae
Amandinea punctata
Buellia disciformis
Buellia dispersa
Buellia erubescens
Buellia triphragmoides
Buellia tyrolensis
Dimelaena oreina
Diplotomma alboatrum
Candelariaceae
Candelaria concolor
Candelariella aurella
Candelariella citrina
Candelariella deflexa
Candelariella rosulans
Candelariella vitellina
Candelariella xanthostigma
Cladoniaceae
Cladonia cariosa
Cladonia chlorophaea
Cladonia coniocraea
Cladonia fimbriata
Cladonia glauca
Cladonia peziziformis
Cladonia pyxidata
Collemataceae
Collema furfuraceum
Collema subflaccidum
Enchylium coccophorum
Enchylium polycarpon
Enchylium tenax
Lathagrium fuscovirens
Scytinium gelatinosum
Graphidaceae
Diploschistes gypsaceus
Diploschistes muscorum
Diploschistes scruposus
Hymeneliaceae
Hymenelia epulotica
Lecanoraceae
Lecanora albellula
Lecanora allophana
Lecanora argopholis
Lecanora cenisia
Lecanora densa
Lecanora nashii
Lecanora neodegelii
Lecanora polytropa
Lecanora rupicola
Lecanora saligna
Lecanora subintricata
Lecanora thallophila
Lecanora valesiaca
Lecanora varia
Lecidella anomaloides
Lecidella carpathica
Lecidella euphorea
Lecidella patavina
Lecidella stigmatea
Lecidella wulfenii
Myriolecis crenulata
Myriolecis dispersa
Myriolecis hagenii
Myriolecis juniperina
Protoparmeliopsis garovaglii
Protoparmeliopsis muralis
Protoparmeliopsis peltata
Rhizoplaca chrysoleuca
Rhizoplaca melanophthalma
Rhizoplaca novomexicana
Rhizoplaca opiniconensis
Rhizoplaca subdiscrepans
Lecideaceae
Lecidea atrobrunnea
Lecidea plana
Lecidea syncarpa
Lecidea tessellata
Porpidia macrocarpa
Lichenotheliaceae
Endococcus oreinae
Lichinaceae
Heppia conchiloba
Heppia lutosa
Lichinella americana
Lichinella nigritella
Lichinella stipatula
Peccania arizonica
Peccania texana
Psorotichia schaereri
Megasporaceae
Aspicilia cinerea
Circinaria caesiocinerea
Circinaria calcarea
Circinaria contorta
Circinaria gibbosa
Lobothallia alphoplaca
Lobothallia praeradiosa
Megaspora verrucosa
Opegraphaceae
Opegrapha pulvinata
Ophioparmaceae
Fulgidea oligospora
Xylopsora friesii
Pannariaceae
Psoroma hypnorum
Vahliella leucophaea
Parmeliaceae
Bryoria fuscescens
Flavopunctelia darrowii
Flavopunctelia soredica
Letharia columbiana
Melanelixia fuliginosa
Melanohalea elegantula
Melanohalea exasperatula
Melanohalea olivacea
Melanohalea subolivacea
Montanelia tominii
Nephromopsis fendleri
Nephromopsis weberi
Protoparmelia badia
Pseudevernia intensa
Usnea cavernosa
Usnea florida
Usnea fulvoreagens
Usnea hirta
Usnea intermedia
Usnea lapponica
Usnea subfloridana
Xanthoparmelia chlorochroa
Xanthoparmelia coloradoensis
Xanthoparmelia cumberlandia
Xanthoparmelia lavicola
Xanthoparmelia lineola
Xanthoparmelia mexicana
Xanthoparmelia plittii
Xanthoparmelia subdecipiens
Xanthoparmelia wyomingica
Peltigeraceae
Nephroma parile
Peltigera canina
Peltigera didactyla
Peltigera horizontalis
Peltigera rufescens
Peltulaceae
Peltula bolanderi
Peltula euploca
Peltula farinosa
Peltula obscurans
Pertusariaceae
Pertusaria saximontana
Phaeococcomycetaceae
Lichenostigma dimelaenae
Phyllachoraceae
Lichenochora verrucicola
Physciaceae
Hyperphyscia adglutinata
Phaeophyscia ciliata
Phaeophyscia endococcina
Phaeophyscia hirsuta
Phaeophyscia nigricans
Phaeophyscia orbicularis
Phaeophyscia sciastra
Physcia adscendens
Physcia aipolia
Physcia albinea
Physcia biziana
Physcia caesia
Physcia callosa
Physcia dubia
Physcia mexicana
Physcia phaea
Physcia stellaris
Physcia tenella
Physconia detersa
Physconia isidiigera
Physconia muscigena
Physconia perisidiosa
Rinodina archaea
Rinodina castanomela
Rinodina coloradiana
Rinodina confragosa
Rinodina juniperina
Rinodina mniaraea
Rinodina zwackhiana
Placynthiaceae
Placynthium nigrum
Psoraceae
Protoblastenia rupestris
Psora cerebriformis
Psora crenata
Psora decipiens
Psora globifera
Psora nipponica
Psora pseudorussellii
Psora russellii
Psora tuckermanii
Psorula rufonigra
Ramalinaceae
Bibbya ruginosa
Lecania brunonis
Lecidea turgidula
Ramalina calicaris
Ramalina fastigiata
Ramalina sinensis
Thalloidima candidum
Thalloidima sedifolium
Toninia tristis
Rhizocarpaceae
Epilichen stellatus
Rhizocarpon arctogenum
Rhizocarpon bolanderi
Rhizocarpon disporum
Rhizocarpon geographicum
Squamarinaceae
Squamarina lentigera
Stereocaulaceae
Lepraria incana
Lepraria neglecta
Teloschistaceae
Amundsenia approximata
Athallia holocarpa
Calogaya decipiens
Calogaya saxicola
Caloplaca allanii
Caloplaca cerina
Caloplaca durietzii
Caloplaca pellodella
Caloplaca pinicola
Caloplaca sideritis
Erichansenia epithallina
Gyalolechia desertorum
Gyalolechia epiphyta
Pachypeltis cladodes
Polycauliona candelaria
Polycauliona polycarpa
Pyrenodesmia atroalba
Pyrenodesmia variabilis
Rufoplaca arenaria
Rusavskia elegans
Solitaria chrysophthalma
Squamulea squamosa
Squamulea subsoluta
Tayloriellina microphyllina
Xanthomendoza fallax
Xanthomendoza montana
Xanthomendoza trachyphylla
Trapeliaceae
Trapeliopsis granulosa
Umbilicariaceae
Umbilicaria hirsuta
Umbilicaria phaea
Verrucariaceae
Catapyrenium rufescens
Catapyrenium squamulosum
Dermatocarpon miniatum
Dermatocarpon moulinsii
Dermatocarpon reticulatum
Endocarpon pusillum
Heteroplacidium compactum
Placidium acarosporoides
Placidium arboreum
Placidium lachneum
Staurothele areolata
Staurothele drummondii
Staurothele effigurata
Staurothele elenkinii
Staurothele monicae
Verrucaria inficiens
Verrucaria muralis