Global occurrence: Eurasia – Europe | Americas – North America (incl Mexico) | Arctic. Substrate: soil, clay, humus, turf, detritus, dead leaves. Life habit: lichenized (mutualistic with algal photobionts). Thallus: squamulose (squamulous); [th] upper surface: grey(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon); [th upper surface]: epruinose | pruinose; [th upper surface] pruina: scarce, sparse, fine; [th] lower surface: black(ish); [th lower surface] specific structures: present. Ascomata: absent | present; ascoma: perithecial, perithecioid – hymenial; ascoma: immersed, innate | subsessile, subimmersed, adnate, semi-immersed, emergent; [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon); [ascm] paraphyses/-oids: absent. Asci: fissitunicate – verrucarialean; [asc] tholus: not thickened | thickened; [asc] tholus amyloidity (iodine reaction): absent. Ascospores: (median) 8.0; [asp] shape: ellipsoidal | ovoid, ovate; [asp] length [µm]: (min) 14.0 (low) 15.0 (high) 18.0 (max) 20.0; [asp] width [µm]: (min) 7.0 (low) 7.5 (high) 9.5 (max) 10.0; [asp] septa: present, transversely septate; [asp] transversal septa: (median) 1.0; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent. Secondary metabolites: absent. Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid. Secondary photobionts (eg in cephalodia): absent.