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Rhizocarpon expallescens Th. Fr.  
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Family: Rhizocarpaceae
[Catocarpus expallescens (Th. Fr.) Hellb.]
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Botanische Staatssammlung Munchen

Global occurrence: Eurasia – Europe | Americas – North America (incl Mexico) | Arctic.  Substrate: rock, stones, pebbles – unspecified.  Life habit: lichenized (mutualistic with algal photobionts).  Thallus: crustose (crustaceous) – episubstratal – unspecified; cracked, fissured, fractured, rimose | compartimentized – areolate, squamulate; [th] upper surface: black(ish) | grey(ish) | grey(ish) white; [th upper surface]: epruinose; [th marginal and upper surface] specific structures: absent; [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (min) 0.5; [th] morphol substructures (eg areoles, squamules): contiguous, coherent (throughout the thallus) | discontiguous, dispersed (throughout the thallus).  Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (min) 1.0; ascoma: sessile, superficial; [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded | subconvex, slightly convex; [ascm, if apoth] disc, mazaedium: black(ish); [ascm, if apoth] disc, mazaedium: epruinose; [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish); [ascm, if apoth] margin excipular photobionts: absent; [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: brown(ish) (if pale: fawn, tan; if mid: cinnamon); [ascm] paraphyses/-oids: present; [ascm] epihymenium, epithecium: green(ish) | blue(ish) black.  Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: with amyloid cap (= Icmadophila-, Lecidea-, Pertusaria-types etc).  Ascospores: (median) 8.0; [asp] shape: ellipsoidal; [asp] length [µm]: (low) 14.0 (high) 17.0; [asp] width [µm]: (low) 6.0 (high) 9.0; [asp] septa: present; [asp] septa: transversely septate; [asp] transversal septa: (median) 1.0; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: thick, distincly halonate.  Secondary metabolites: absent.  Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid.  Secondary photobionts (eg in cephalodia): absent.

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This project made possible by National Science Foundation Awards: #1115116