Global occurrence: Africa (incl Madagascar) | Eurasia – Asia Extratropical | Oceania – Australasia | Eurasia – Europe | Americas – North America (incl Mexico) | Americas – South and Central America | Arctic. Substrate: rock – calcareous, calciferous, basic | rock, stones, pebbles – unspecified. Life habit: lichenized (mutualistic with algal photobionts). Thallus: crustose (crustaceous) – episubstratal – unspecified; cracked, fissured, fractured, rimose | compartimentized – areolate, squamulate; [th] upper surface: grey(ish) | white(ish); [th upper surface]: pruinose; [th marginal and upper surface] specific structures: absent. Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma: immersed, innate | subsessile, subimmersed, adnate, semi-immersed, emergent; [ascm] paraphyses/-oids: present. Asci: unitunicate; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): absent. Ascospores: (median) 4.0; [asp] shape: broadly ellipsoidal; [asp] length [µm]: (min) 22.0 (low) 30.0 (high) 40.0 (max) 45.0; [asp] width [µm]: (min) 10.0 (low) 15.0 (high) 18.0 (max) 20.0; [asp] septa: present, muriformly septate – muriform, submuriform; [asp] transversal septa: (low) 4.0 (high) 7.0; [asp] pigmentation: medium brown, brownish; [asp] perispore, epispore: not apparent. Secondary metabolites: present, lecanoric acid. Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid. Secondary photobionts (eg in cephalodia): absent.