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Bellemerea cupreoatra (Nyl.) Clauzade & Cl. Roux  
Family: Lecideaceae
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Botanische Staatssammlung Munchen

Global occurrence: Eurasia – Asia Extratropical | Eurasia – Europe.  Substrate: rock, stones, pebbles – unspecified.  Life habit: lichenized (mutualistic with algal photobionts).  Thallus: crustose (crustaceous) – episubstratal – unspecified; thallus: compartimentized – areolate, squamulate; [th] upper surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon); [th marginal and upper surface] specific structures: absent; [th] medulla amyloidity: absent | present.  Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma: immersed, innate; [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded; [ascm, if apoth] disc, mazaedium: brown(ish) black; [ascm, if apoth] disc, mazaedium: epruinose; [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: hyaline, colourless; [ascm] paraphyses/-oids: present; [ascm] epihymenium, epithecium: brown(ish) (if pale: fawn, tan; if mid: cinnamon).  Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: with amyloid tube, ring (= Collema-, Micarea-, Porpidia-, Psora-types etc).  Ascospores: (median) 8.0; [asp] shape: ellipsoidal; [asp] length [µm]: (low) 5.0 (high) 10.0; [asp] width [µm]: (low) 5.0 (high) 9.0; [asp] septa: absent – spore lumen unilocular, monolocular; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent | thin, thinly halonate | thick, distincly halonate.  Secondary metabolites: present, gyrophoric acid.  Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid.  Secondary photobionts (eg in cephalodia): absent.

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This project made possible by National Science Foundation Awards: #1115116