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Hypogymnia metaphysodes (Asah.) Rass.  
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Family: Parmeliaceae
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Stephen Sharnoff  
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Botanische Staatssammlung Munchen

Global occurrence: Americas – North America (incl Mexico) | Arctic.  Substrate: wood – dead, living | bark, cork, plant surface – trunks, branches, twigs.  Life habit: lichenized (mutualistic with algal photobionts).  Thallus: foliose (foliaceous), leaf-like; [th] upper surface: grey(ish) | green(ish) grey; [th upper surface]: epruinose; [th marginal and upper surface] specific structures: absent; [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 1.0 (high) 2.0; [th] morphol substructures (eg areoles, squamules): distantly discontiguous; [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane; [th] morphol substructures (eg lobes, branches): subdichotomous; [th] lower surface: black(ish); [th lower surface] specific structures: absent.  Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma: subpedicellate, substipitate, subpedunculate, substalked; [ascm, if apoth] disc, mazaedium: concave; [ascm, if apoth] disc, mazaedium: yellow(ish) brown; [ascm, if apoth] disc, mazaedium: epruinose; [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: grey(ish) | green(ish) grey; [ascm, if apoth] margin excipular photobionts: present | abundant; [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: hyaline, colourless; [ascm] paraphyses/-oids: present; [ascm] epihymenium, epithecium: yellow(ish) brown | red(dish) brown (pale: orange brown).  Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: amyloid with widening axial body towards the apex (= Lecanora-, Parmelia-, Rinodina-types etc).  Ascospores: (median) 8.0; [asp] shape: ovoid, ovate; [asp] length [µm]: (median) 6.0; [asp] width [µm]: (median) 4.0; [asp] septa: absent – spore lumen unilocular, monolocular; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent.  Secondary metabolites: present, atranorin | chloroatranorin | physodic acid | 3-hydroxyphysodic acid.  Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid.  Secondary photobionts (eg in cephalodia): absent.

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Stephen Sharnoff  
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Stephen Sharnoff  
Hypogymnia metaphysodes image
Stephen Sharnoff  
Hypogymnia metaphysodes image
Stephen Sharnoff  
Hypogymnia metaphysodes image
Hypogymnia metaphysodes image
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This project made possible by National Science Foundation Awards: #1115116, #2001500, #2001394