Global occurrence: Eurasia – Asia Extratropical | Eurasia – Europe | Arctic. Substrate: lichens, lichenized fungi | rock – siliceous, siliciferous, acidic | rock, stones, pebbles – unspecified. Life habit: lichenized (mutualistic with algal photobionts) | lichenicolous (in/on lichens). Thallus: crustose (crustaceous) – episubstratal – unspecified | subsquamulose (subsquamulous); thallus: compartimentized – areolate, squamulate; [th] upper surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | grey(ish) brown| grey(ish) | blue(ish) grey | brown(ish) grey; [th upper surface]: epruinose; [th marginal and upper surface] specific structures: absent | present; [th margin] cilia, cilioid structures: absent; [th upper surface] isidia, isidioid structures: absent; [th upper surface] soredia, soralia, soralioid structures: present; [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 0.2 (low) 0.4 (high) 1.0 (max) 1.8; [th] morphol substructures (eg areoles, squamules): discontiguous, dispersed (throughout the thallus); [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane; [th] medulla amyloidity: absent. Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (min) 0.2 (low) 0.5 (high) 0.7 (max) 1.1; ascoma: immersed, innate | subsessile, subimmersed, adnate, semi-immersed, emergent; [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded | subconvex, slightly convex; [ascm, if apoth] disc, mazaedium: black(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) | black(ish) brown | red(dish) brown (if pale: orange brown); [ascm, if apoth] disc, mazaedium: epruinose; [ascm, if apoth] margin excipular photobionts: absent; [ascm, if apoth] subhymenial layers, hypothecium, [if perith] basal excipulum: hyaline, colourless; [ascm] paraphyses/-oids: present; [ascm] paraphyses/-oid cells width [µm]: (low) 1.5 (high) 2.0; [ascm] paraphyses/-oid apical cells width [µm]: (min) 2.5 (low) 3.0 (high) 4.5; [ascm] epihymenium, epithecium: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | yellow(ish) brown | red(dish) brown (pale: orange brown). Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: amyloid with widening axial body towards the apex (= Lecanora-, Parmelia-, Rinodina-types etc). Ascospores: (median) 8.0; [asp] shape: ellipsoidal | elongate-fusiform, ellipsoidal-fusiform; [asp] length [µm]: (min) 8.0 (median) 10.5 (high) 16.0 (max) 18.0; [asp] width [µm]: (min) 4.0 (low) 5.0 (high) 6.0 (max) 7.0; [asp] septa: absent – spore lumen unilocular, monolocular; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent. Secondary metabolites: present, miriquidic acid. Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid. Secondary photobionts (eg in cephalodia): absent.