Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2002. Lichen Flora of the Greater Sonoran Desert Region. Vol 1.

Thallus: continuous, immersed in the substrate upper surface: whitish, smooth dull photobiont: absent Perithecia: hemispherical, black, semi-immersed in the thallus, c. 0.4-1 mm diam. involucrellum: dark brown; exciple thin, pale brown ascomatal wall: black, absent below the hamathecium hamathecium: branched pseudoparaphyses, anastomosing above the asci; filaments: c. 1 micrometer wide, not inspersed with oil droplets asci: cylindrical to clavate, c. 70-90 x 20-23 micrometer, thick-walled, with 2-6, irregularly arranged ascospores ascospores: hyaline, long ellipsoid to clavate, densely muriform, 22-28 x 9-13 micrometer; walls: not ornamented, with a c. 2-4 micrometer wide gelatinous sheath Pycnidia: not seen Spot tests: all negative, UV negative Secondary metabolites: none detected. Substrate and ecology: on bark of various woody plants, including Euphorbia and Rhus World distribution: pantropical or possibly nearly cosmopolitan Sonoran distribution: collected several times in California (including the Channel islands), Baja California and in Sonora.