Ecology: Terricolous or bryophytic (over rocks generally in or around bogs); growing sun-exposed or partially shaded. Thallus: Crustose, squamulose; lobes ascending or horizontal; branches loosely aggregated, clustered, or densely aggregated; separate thallus parts 1-4 mm long; .5-1.5 mm wide. Primary Thallus: Evanescent or persistent. Secondary Thallus: Present, of indeterminate growth; stipe 15-70 mm high, .6-3 mm wide, lime green, olive, or brown (121,95,79,57-55, browner in s. str., greener in "japonica"), mottled or tainted brown, corticate, with intact or rarely longitudinally split wall, 20-40 µm thick. Thallus Outline: Margin crenulate or dissected. Upper Surface: Areolated (paler patches on a dark brown to black background), lime green, olive, or brown (121,95,79, 57-55), smooth; not sorediate. Lower Surface: White or brown (at base). Lower Cortex: Absent. Ascocarps: Forming along the margin of scyphi, moderately abundant, .15-.8 mm in diam.. Disk: Plane (emarginate, or with a thin margin even with the disk); brown (greyish red-brown (46-47)). Conidiomata: Formed on margins of scyphi (abundant). Pycnidia: With reddish ostiolar gel. Secondary Metabolites: Chemotype 1: Squamatic acid, of the following substance class(es): ß-orcinol depsides. Chemotype 2: Thamnolic acid (often with traces of decarboxythamnolic acid) or barbatic acid (traces), of the following substance class(es): ß-orcinol depsides. Spot Tests: Chemotype 1: Upper surface: K (negative), C , PD . Chemotype 2: Upper surface: K + deep yellow, C , PD + orange. UV-Fluorescence: Reaction colour: bright bluish white.