Global occurrence: Eurasia – Europe | Americas – North America (incl Mexico) | Arctic. Substrate: bryophytes – mosses, liverworts. Life habit: lichenized (mutualistic with algal photobionts). Thallus: foliose (foliaceous), leaf-like; continuous, diffuse, effuse; [th] upper surface: grey(ish) | grey(ish) brown; [th upper surface]: epruinose; [th marginal and upper surface] specific structures: absent; [th] morphol substructures (eg areoles, lobes, branches) upper surface: verruculose, minutely warted; [th] lower surface: brown(ish) yellow (ochraceous, ochre); [th lower surface] specific structures: present; [th lower surface] rhizines, rhizoid structures: present; [th lower surface] rhizines, rhizinoid structures: unbranched, simple | fasciculate. Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; [ascm] paraphyses/-oids: present. Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: with amyloid tube, ring (= Collema-, Micarea-, Porpidia-, Psora-types etc). Ascospores: (median) 8.0; [asp] length [µm]: (low) 80.0 (high) 95.0; [asp] width [µm]: (low) 3.5 (high) 4.0; [asp] septa: present, transversely septate; [asp] transversal septa: (median) 3.0; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent. Secondary metabolites: present; 15α-acetoxyhopane-22-ol (dolichorrhizin) | methyl gyrophorate | tenuiorin | zeorin (hopane-6α,22-diol) | unknown/unidentified (tri-)terpenoid(s), (tri-)terpene(s). Primary photobiont: present, cyanobacterial. Secondary photobionts (eg in cephalodia): absent.