Life habit: lichenized (mutualistic with algal photobionts) thallus: squamulose (squamulous) | subfruticose (subfruticous), fruticulose, dactyliform thallus: compartimentized – areolate, squamulate [th] upper surface: black(ish) | yellow(ish) green | white(ish) green | green(ish) yellow | white(ish) red (rose) [th upper surface]: epruinose | pruinose [th upper surface] pruina: scarce, sparse, fine | dense [th marginal and upper surface] specific structures: absent [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 1.5 (high) 2.0 [th] morphol substructures (eg areoles, squamules): distantly discontiguous [th] morphol substructures (eg areoles, lobes, branches) upper surface: subsquamulose | squamulose [th lower surface] specific structures: absent | present [th lower surface] rhizines, rhizoid structures: absent ascomata: absent | present ascoma: apothecial, apothecioid – hymenial ascoma [mm]: (low) 3.0 (high) 4.0 ascoma: sessile, superficial [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded [ascm, if apoth] disc, mazaedium: black(ish) | blue(ish) black | brown(ish) black | green(ish) grey | grey(ish) brown | yellow(ish) brown [ascm, if apoth] disc, mazaedium: pruinose [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish) | yellow(ish) green | white(ish) green | green(ish) yellow | white(ish) red (rose) | red(dish) white (pink) [ascm, if apoth] margin excipular photobionts: present | abundant [ascm] paraphyses/-oids: present [ascm] paraphyses/-oid cells width [µm]: (low) 2.0 (high) 2.5 [ascm] paraphyses/-oid apical cells width [µm]: (low) 2.5 (high) 3.0 [ascm] epihymenium, epithecium: brown(ish) (if pale: fawn, tan; if mid: cinnamon) asci: lecanoralean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): present [asc] tholus amyloidity pattern: amyloid with widening axial body towards the apex (= Lecanora-, Parmelia-, Rinodina-types etc) ascospores: (median) 8.0 [asp] shape: ellipsoidal | elongate-fusiform, ellipsoidal-fusiform | ovoid, ovate [asp] length [µm]: (low) 11.0 (high) 15.0 (max) 19.0 [asp] width [µm]: (low) 6.0 (high) 8.0 (max) 9.0 [asp] septa: absent – spore lumen unilocular, monolocular [asp] pigmentation: hyaline, colourless [asp] perispore, epispore: not apparent conidiomata: absent | present conidia: [cur] – curved, unciform, falcate, sickle-shaped [co] length [µm]: (low) 11.0 (high) 17.0 (max) 20.0 secondary metabolites: present secondary metabolites: 16ß-acetoxyhopane-6α,22-diol | 6α,16β-diacetoxyhopane-22-ol | 6α-acetoxyhopane-16β,22-diol | hopane-6α,16ß,22-triol | isousnic acid | usnic acid | zeorin (hopane-6α,22-diol) primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid

substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts.