Global occurrence: Eurasia – Europe | Americas – North America (incl Mexico) | Arctic. Substrate: lichens, lichenized fungi. Life habit: lichenized (mutualistic with algal photobionts) | lichenicolous (in/on lichens). Thallus: crustose (crustaceous) – episubstratal – unspecified; compartimentized – areolate, squamulate; [th] upper surface: grey(ish) | white(ish) grey; [th marginal and upper surface] specific structures: absent; [th] medulla amyloidity: present. Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (low) 0.25 (high) 0.8; ascoma: immersed, innate | subsessile, subimmersed, adnate, semi-immersed, emergent | sessile, superficial; [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded | subconvex, slightly convex | convex; [ascm, if apoth] disc, mazaedium: black(ish); [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish); [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: brown(ish) (if pale: fawn, tan; if mid: cinnamon); [ascm] paraphyses/-oids: present; [ascm] epihymenium, epithecium: black(ish) green | brown(ish) green (olivaceous, olive green) | green(ish) brown (olive brown). Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: amyloid with acuate axial body towards the apex (= Bacidia-, Buellia-, Ramalina-types etc). Ascospores: (median) 8.0; [asp] shape: oblong, oblong-obtuse; [asp] length [µm]: (min) 6.0 (low) 8.0 (high) 10.0 (max) 11.2; [asp] width [µm]: (min) 3.5 (low) 5.3(median) 5.7 (high) 6.0; [asp] septa: present, transversely septate; [asp] transversal septa: (median) 1.0; [asp] pigmentation: medium brown, brownish; [asp] perispore, epispore: ornamented, rough, warted, striate, cracked. Conidiomata: absent | present. Conidia: bacilliform, bacillary; [co] length [µm]: (low) 3.5 (high) 4.0. Secondary metabolites: present, miriquidic acid. Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid. Secondary photobionts (eg in cephalodia): absent.