Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2004. Lichen Flora of the Greater Sonoran Desert Region. Vol 2.
Thallus: crustose, cracked to areolate, thick, rugose surface: cream-colored, pale yellow or gray dull, roughened, with small crystals Apothecia: embedded in the thallus or prominently raised together with the surrounding thallus, lirellate lirellae: oblong, slightly flexuous, rarely branched, 1-8 x 0.5-1 mm disc: slit-like, not visible margin: often whitish gray, only the slit gray or dark brown; excipular lips: striate, slightly crenate, dark brown but almost completely covered by a ±thin, whitish thalline cover exciple: well developed and completely carbonized, basal part thick, up to 300 µm high and often with many large crystals; excipular lips: carbonized, striate, but sometimes only apices distinctly striate, with ±complete thalline cover epihymenium: brown, 3-5 µm high, sometimes poorly developed hymenium: hyaline, 160-200 µm tall, upper part fine inspersed (in the Sonoran sample); paraphyses asci: clavate, 120-140 x 15-25 µm (±immature; the shape of the mature ascus is quasi identical to the ascospore size), 1-spored ascospores: hyaline, muriform, 22-27 x 7-9-locular, 100-160 x 35-60 µm, I+ dark violet Pycnidia and conidia: not found Spot tests: cortex and medulla K-, C-, KC-, P- Secondary metabolites: none detected. Substrate and ecology: on sandstone World distribution: tropical to subtropical, so far reported from Brazil, Hawaii, and Mexico Sonoran distribution: Sierra Madre Occidental region of Chihuahua. Notes: Previously Graphis pseudosophistica was only known on bark (rarely on mosses). The cited collection from the Sonoran region corresponds morphologically well with the type, but grows on rock and shows a fine inspersion of the upper part of the hymenium. As little is known about the substrate specifity of this species, we conclude that the rock sample from Mexico is Graphis pseudosophistica.