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Family:
Umbilicariaceae
[Actinogyra rigida (Du Rietz) Llano, more, Agyrophora rigida (Du Rietz) Llano, Gyrophora anthracina (Wulfen) Körb., Gyrophora rigida Du Rietz, Gyrophora rigida var. discissa (Th. Fr.) Du Rietz, Gyrophora rigida var. rigida Du Rietz, Umbilicaria coriacea Imshaug]
 Stephen Sharnoff |
Life habit: lichenized (mutualistic with algal photobionts) thallus: peltate, umbilicate thallus: continuous, diffuse, effuse [th] upper surface: grey(ish) | white(ish) | brown(ish) black [th upper surface]: pruinose [th upper surface] pruina: dense [th marginal and upper surface] specific structures: absent | present [th margin] cilia, cilioid structures: absent [th upper surface] isidia, isidioid structures: absent [th upper surface] soredia, soralia, soralioid structures: absent [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 20.0 (high) 50.0 [th] morphol substructures (eg areoles, squamules): distantly discontiguous [th] morphol substructures (eg areoles, lobes, branches) upper surface: verrucose, warted | rugose, plicate, folded, pustulate, faveolate, wrinkled [th] lower surface: grey(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) [th lower surface] specific structures: present [th lower surface] rhizines, rhizoid structures: absent ascomata: absent | present ascoma: apothecial, apothecioid – hymenial ascoma [mm]: (low) 1.0 (high) 2.0 ascoma: subpedicellate, substipitate, subpedunculate, substalked | pedicellate, stipitate, pedunculate, stalked [ascm, if apoth] disc, mazaedium: concave | plane, flat, flattened, expanded [ascm, if apoth] disc, mazaedium: black(ish) [ascm, if apoth] disc, mazaedium: epruinose [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish) [ascm, if apoth] margin excipular photobionts: absent [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: brown(ish) (if pale: fawn, tan; if mid: cinnamon) [ascm] paraphyses/-oids: present [ascm] paraphyses/-oid cells width [µm]: (low) 2.0 (high) 2.5 [ascm] paraphyses/-oid apical cells width [µm]: (low) 2.5 (high) 4.0 [ascm] epihymenium, epithecium: black(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) asci: lecanoralean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): present [asc] tholus amyloidity pattern: entirely amyloid (= Catillaria-, Teloschistes-types etc) ascospores: (median) 8.0 [asp] shape: ellipsoidal [asp] length [µm]: (low) 9.5 (high) 14.5 [asp] width [µm]: (low) 2.5 (high) 9.0 [asp] septa: absent – spore lumen unilocular, monolocular [asp] pigmentation: hyaline, colourless [asp] perispore, epispore: not apparent conidiomata: absent | present conidia: cylindrical [co] length [µm]: (low) 3.3 (high) 4.0 secondary metabolites: present secondary metabolites: norstictic acid primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid. substrate: rock – siliceous, siliciferous, acidic.
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