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Umbilicaria havaasii Llano
Family: Umbilicariaceae
Umbilicaria havaasii image
Lucy Taylor  
life habit: lichenized (mutualistic with algal photobionts) thallus: peltate, umbilicate thallus: continuous, diffuse, effuse [th] upper surface: black(ish) | grey(ish) [th upper surface]: pruinose [th upper surface] pruina: scarce, sparse, fine | dense [th marginal and upper surface] specific structures: absent [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 80.0 (high) 130.0 [th] morphol substructures (eg areoles, squamules): distantly discontiguous [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane | granulose, granular | rugose, plicate, folded, pustulate, faveolate, wrinkled [th] lower surface: black(ish) | red(dish) brown (if pale: orange brown) [th lower surface] specific structures: absent | present [th lower surface] rhizines, rhizoid structures: absent | present [th lower surface] rhizines, rhizinoid structures: unbranched, simple | furcately branched, furcate, forked ascomata: absent | present ascoma: apothecial, apothecioid – hymenial ascoma [mm]: (low) 0.9 (high) 2.0 ascoma: sessile, superficial [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded | subconvex, slightly convex [ascm, if apoth] disc, mazaedium: black(ish) [ascm, if apoth] disc, mazaedium: epruinose [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish) [ascm, if apoth] margin excipular photobionts: absent [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: brown(ish) (if pale: fawn, tan; if mid: cinnamon) [ascm] paraphyses/-oids: present [ascm] paraphyses/-oid cells width [µm]: (low) 1.7 (high) 2.0 [ascm] paraphyses/-oid apical cells width [µm]: (low) 7.0 (high) 9.5 [ascm] epihymenium, epithecium: black(ish) asci: lecanoralean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): present [asc] tholus amyloidity pattern: entirely amyloid (= Catillaria-, Teloschistes-types etc) ascospores: (median) 8.0 [asp] shape: ellipsoidal [asp] length [µm]: (low) 10.0 (high) 20.5 [asp] width [µm]: (low) 6.0 (high) 8.5 [asp] septa: absent – spore lumen unilocular, monolocular [asp] pigmentation: hyaline, colourless [asp] perispore, epispore: not apparent secondary metabolites: present secondary metabolites: gyrophoric acid primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: rock – siliceous, siliciferous, acidic.