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Protoblastenia incrustans (DC.) J. Steiner
Family: Psoraceae
[Biatora incrustans (DC.) A. Massal.,  more]
Protoblastenia incrustans image
André Aptroot  

Global occurrence: Eurasia – Asia Extratropical | Eurasia – Europe | Americas – North America (incl Mexico) | Arctic. Substrate: rock – calcareous, calciferous, basic | rock, stones, pebbles – unspecified.  Life habit: lichenized (mutualistic with algal photobionts).  Thallus: crustose (crustaceous) – endosubstratal, inconspicuous, immersed | crustose (crustaceous) – episubstratal – unspecified; continuous, diffuse, effuse | cracked, fissured, fractured, rimose; [th] upper surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | white(ish) | white(ish) grey; [th marginal and upper surface] specific structures: absent.  Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (min) 0.2 (low) 0.3(median) 0.4 (high) 0.55 (max) 0.85; ascoma: immersed, innate | subsessile, subimmersed, adnate, semi-immersed, emergent; [ascm, if apoth] disc, mazaedium: concave | plane, flat, flattened, expanded | subconvex, slightly convex; [ascm, if apoth] disc, mazaedium: red(dish) yellow (orange, orange yellow) | yellow(ish) red (orange red); [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: yellow(ish) red (orange red); [ascm, if apoth] margin excipular photobionts: absent; [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: yellow(ish) (if dull: buff) | hyaline, colourless; [ascm] paraphyses/-oids: present; [ascm] paraphyses/-oid cells width [µm]: (low) 1.5(median) 2.5 (high) 3.5; [ascm] paraphyses/-oid apical cells width [µm]: (low) 2.0(median) 3.2 (high) 4.5; [ascm] epihymenium, epithecium: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | red(dish) yellow (orange, orange yellow) | yellow(ish) red (orange red).  Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: with amyloid tube, ring (= Collema-, Micarea-, Porpidia-, Psora-types etc).  Ascospores: (median) 8.0; [asp] shape: broadly ellipsoidal | ellipsoidal | ovoid, ovate; [asp] length [µm]: (low) 9.0(median) 10.9 (high) 12.5 (max) 13.0; [asp] width [µm]: (min) 4.0 (low) 4.5(median) 5.6 (high) 6.0 (max) 7.0; [asp] septa: absent – spore lumen unilocular, monolocular; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent.  Conidiomata: absent | present.  Conidia: cylindrical | bacilliform, bacillary.  Secondary metabolites: present, unknown/unidentified anthraquinone(s).  Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid.  Secondary photobionts (eg in cephalodia): absent.