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Family:
Physciaceae
[Berengeria confragosa (Ach.) Trevis., more, Lecanora confragosa (Ach.) Röhl., Lecanora confragosa var. confragosa (Ach.) Röhl., Lecanora confragosa var. lecidotropa Nyl., Parmelia confragosa Ach., Rinodina exigua var. confragosa (Ach.) Th. Fr., Rinodina metabolica var. confragosa (Ach.) Hazsl., Zeora confragosa (Ach.) Flot.]
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Thallus: crustose, usually thin, consisting of more or less discrete areoles, up to c. 0.5 mm wide sometimes with minutely lobate margins, or thick, with contiguous areoles, up to 0.5-0.7 mm wide, plane or rugose surface: light gray, dull; margin: indeterminate, rarely determinate; prothallus: present on smooth surfaces, dark, fimbriate; vegetative propagules: absent Apothecia: sessile, frequent, typically scattered, up to 0.7-1 mm in diam. disc: black, plane becoming slightly convex thalline margin: concolorous with thallus, entire, c. 0.1 mm wide, persistent or becoming partly excluded; excipular ring: sometimes present, raised thalline exciple: 80-90 µm wide laterally; cortex: 10-20 µm wide; cells: up to c. 5 µm wide, not pigmented; algal cells: up to 12-17 µm in diam.; thalline exciple: 100120 µm thick below; cortex: 30-50 µm thick, columnar proper exciple: hyaline, 5-10 µm wide laterally, expanding to 15-25 µm at periphery hymenium: 80-90 µm tall; paraphyses: 2-3 µm wide, not conglutinate, with apices up to 3.5-5 µm, lightly pigmented, immersed in a dispersed pigment forming a red-brown epihymenium; hypothecium: hyaline, 40-70 µm thick asci: clavate, 50-60 x 16-17 µm, 8-spored ascospores: brown, 1-septate, ellipsoid, developmental type A, Physcia-type, (16.5-)19-20.5(-22.5) x (8-)9-10(-11) µm; lumina becoming inflated but retaining thick apical wall; torus: pigmented; walls: ornamented (Fig. 62) Pycnidia: immersed in thallus, ostioles pigmented; conidiophores: Type I conidia: bacilliform, 3.5-5.5 x 1-1.5 µm Spot tests: K+ yellow, C-, KC-, P- or P+ faint yellow Secondary metabolites: atranorin in cortex, zeorin in medulla. Substrate and ecology: a shade tolerant species of siliceous and basaltic rocks, frequently being found on vertical or overhanging rock faces, and boulder fields in montane habitats World distribution: Europe, Africa and western North America (Rocky Mountains from Alberta to Arizona, Black Hills, coastal ranges, Sierra Nevada) Sonoran distribution: mountains of Arizona, coastal ranges of southern California and Baja California, at elevations of 1375-3540 m. Notes: Rinodina confragosa is in many ways the saxicolous equivalent of R. capensis, possessing Physcia-type spores, a columnar lower cortex, and atranorin and zeorin. The species is similar in thallus morphology to some forms of R. bolanderi but that species has larger, Teichophilatype spores.
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