Life habit: lichenized (mutualistic with algal photobionts) thallus: squamulose (squamulous) | subsquamulose (subsquamulous) [th] upper surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | brown(ish) yellow (ochraceous, ochre) [th marginal and upper surface] specific structures: absent [th] lower surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon) [th lower surface] specific structures: present [th lower surface] rhizines, rhizoid structures: present ascomata: absent | present ascoma: apothecial, apothecioid – hymenial ascoma: immersed, innate | subsessile, subimmersed, adnate, semi-immersed, emergent | sessile, superficial [ascm] paraphyses/-oids: present [ascm] epihymenium, epithecium: brown(ish) (if pale: fawn, tan; if mid: cinnamon) asci: lecanoralean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): present [asc] tholus amyloidity pattern: amyloid with widening axial body towards the apex (= Lecanora-, Parmelia-, Rinodina-types etc) ascospores: (median) 8.0 [asp] length [µm]: (low) 18.0 (high) 22.0 [asp] width [µm]: (low) 8.0 (high) 10.0 [asp] septa: present [asp] septa: transversely septate [asp] transversal septa: (median) 1.0 [asp] pigmentation: pale brown | medium brown, brownish | dark brown secondary metabolites: present secondary metabolites: variolaric acid | unknown/unidentified (tri-)terpenoid(s), (tri-)terpene(s) primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts.