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Family:
Parmeliaceae
[Parmelia conspersa f. lacinulata (Gyeln.) Gyeln., more, Parmelia conspersa var. incisa (Taylor) Zahlbr., Parmelia conspersa var. laxa Müll. Arg., Parmelia incisa Taylor, Parmelia laxa (Müll. Arg.) Gyeln., Parmelia laxa f. balatonica (Gyeln.) Gyeln., Parmelia laxa f. borealis Gyeln., Parmelia laxa f. caespitosa (Müll. Arg.) Gyeln., Parmelia laxa f. dentaticola (Gyeln.) Gyeln., Parmelia laxa f. felkaensis (Gyeln.) Gyeln., Parmelia laxa f. lacinulata Gyeln., Parmelia laxa f. laxa (Müll. Arg.) Gyeln., Parmelia laxa f. montevidensis (Müll. Arg.) Gyeln., Parmelia laxa f. parvala Gyeln., Parmelia laxa f. parvula Gyeln., Parmelia laxa f. pueblana Gyeln., Parmelia laxa f. strigosa (Müll. Arg.) Gyeln., Parmelia laxa f. subtananarivensis Gyeln., Parmelia laxa var. laxa (Müll. Arg.) Gyeln., Parmelia laxa var. mediterranea (Gyeln.) Gyeln., Parmelia laxa var. pulvinaris (Zahlbr.) Gyeln., Parmelia laxa var. rosettiformis Gyeln., Parmelia laxa var. subdiffluens Zahlbr. ex Gyeln., Parmelia laxa var. terricola (Gyeln.) Gyeln., Parmelia pulvinaris var. lacinulata Gyeln., Parmelia stramineonitens Zahlbr., Parmelia tasmanica Tayl., Squamarina incisa (Fr.) C.W. Dodge]
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Thallus: foliose, loosely adnate, 6-15 cm in diam., irregularly lobate lobes: subirregular, elongate, plane to subconvex, separate and often forming an imbricate mat, 2-5 mm wide, not lobulate; apices: subrotund, smooth to crenate, eciliate upper surface: yellow to yellow-green, smooth but becoming strongly rugose with age, dull to slightly shiny, epruinose and emaculate, without soralia, isidia, or pustulae medulla: white, with continuous algal layer lower surface: black, plane, shiny, sparsely rhizinate; rhizines: black, simple, 0.3-1 mm long Apothecia: common, substipitate, 2-15 mm wide, laminal on thallus; disc: cinnamon-brown to dark brown; margin: smooth, pruina absent asci: clavate, 8-spored ascospores: hyaline, simple, ellipsoid, 9-11 x 5-7 µm Pycnidia: common, immersed conidia: bifusiform, 5-7 x 0.5 µm Spot tests: upper cortex K-, C-, KC-, P-; medulla K+ yellow becoming dark red, C, KC-, P+ red Secondary metabolites: upper cortex with usnic acid (major); medulla with salazinic acid (major) and consalazinic acid (minor) and often with norstictic and protocetraric acids (both trace). Substrate and ecology: on acidic rocks, often in open or partially shaded habitats World distribution: Australasia, Africa, North and South America Sonoran distribution: occasional at intermediate elevations in Arizona, southern California, Baja California and Chihuahua. Notes: This widely distributed species is characterized by its large, conspicuous thalli with a jet black lower surface, the lack of maculae and vegetative propagules and the presence of salazinic acid in the medulla. Approximately the same range of morphological variation was observed in the Sonoran populations as in the Australasian ones. In the Sonoran region it could be confused with X. subtasmanica, the distinguishing features of which are described above under that species.
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