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Family:
Lecideaceae
[Biatora geophana (Nyl.) Th. Fr., more, Biatorella geophana (Nyl.) Rehm, Biatorella geophana f. boreella (Nyl.) Vain., Biatorella geophana f. geophana (Nyl.) Rehm, Lecidea geophana Nyl., Lecidea geophana var. geophana Nyl., Lecidea geophana var. mucosa Erichsen, Lecidea insita Stirt., Nesolechia insita (Stirt.) Vouaux, Pleolecis geophana (Nyl.) Clem., Sarcogyne geophana (Nyl.) Boistel]
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Life habit: lichenized, ephemeral Thallus: crustose, thin, inconspicuous, membranous, effuse; prothallus: absent surface: pale gray to dull olive, rarely cracked, lacking asexual reproductive structures photobiont: primary one a chlorococcoid green alga, secondary one absent Apothecia: scattered, partially immersed to sessile, 0.2-0.6 mm in diam., lacking a thalline exciple disc: dark brownish black, strongly convex proper exciple: poorly developed, consisting of undifferentiated hyphae epihymenium: red-brown to brown, with irregular granules not dissolving in K hymenium: hyaline to partially pale red-brown, 65-75 µm tall; paraphyses: short, scarce, c. 0.7 µm wide, without expanded apices; subhymenium: pale to dark brown, composed of vertically oriented hyphae; hypothecium: yellowish brown asci: clavate-cylindrical, thin-walled, without a tholus, K/I + blue, with a distinct apical pore, (12-)16-spored ascospores: simple, hyaline, globose to very broadly ellipsoid, 5-7 µm in diam.; wall: thick, c. 0.5 µm wide Conidiomata: pycnidial, black; conidiogenous cells: elongate-ampulliform, enteroblastic, acrogenous conidia: simple, hyaline, ellipsoid, 1-1.5 x 0.5 µm Spot tests: thallus K-, C-, KC-, P- Secondary metabolites: none detected. Substrate: primarily on disturbed soil that is often metal rich, occasionally on wood, bones or man-made materials Geography: Europe and North America Sonoran distribution: high mountains of Arizona. Notes: Steinia geophana is easily overlooked because of its small size and ephemeral nature. It is characterized by a poorly developed true exciple and multispored asci with globose spores
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