Ecology: Biotroph; lichenized; episubstratic; substrate non-calciferous. Thallus: Not subdivided parts, areolate (primarily areolate), continuous throughout to dispersed along the periphery, continuous in the centre; lobes angular at the front; separate thallus parts.2-1 mm thick. Thallus Size and Differentiation: Large; main branches basally not constricted. Thallus Outline: Margin concolorous the thallus centre. Upper Surface: Lemon (citrine) to grey to brown (grey-beige), plane to convex; not sorediate; without thalloconidia thalloconidia. Medulla: Iodine reaction in Lugol's solution positive. Ascocarps: Forming on the pro- or hypothallus, rarely soon sessile, not constricted to slightly constricted at the base, 2-70 per cm², .9-1.6-(4.5) mm in diam.. Margin: Not flexuose, persistent to excluded, 35-80 µm wide, epruinose. Disk: Plane to weakly convex to strongly convex, black, epruinose, pruina scarce, without an umbo. Exciple: Hyphae radiating outwards to intricately interwoven, with no predominant orientation; 2.5-3.5 µm wide; grey; inspersed with crystals. Epithecium: Apical cells olive, cell pigment reacting with 10% HCl (in H2O). Hymenium: 50-70 µm high; white; iodine reaction: Lugol’s positive. Subhymenium: 19-45 µm high; white. Hypothecium: Brownish yellow to brown. Ascospores: Ellipsoid to oblong, (6.5)-9-13.9-(18) µm long, (3.5)-4.1-6.7-(8) µm wide; wall not ornamented. Conidia: (7)-9.2-13.5-(18) µm long. Secondary Metabolites: Present, confluentic acid to 2'-O-methylmicrophyllinic acid to 2'-O-methylperlatolic acid to stictic acid. Spot Tests: Medulla: K –, C –, PD – ascocarp margin (in section): K –, C – disk: C – hypothecium: K – epihymenium: K –.