: Ecology: Substrate non-calciferous. Thallus: Rimose or areolate (primarily areolate), bullate; separate thallus parts thick. Thallus Outline: Indistinct; soon disappearing; white. Upper Surface: Verrucose or verruculose, lemon (citrine), matt, smooth; epruinose; not sorediate. Ascocarps: Strongly constricted at the base. Margin: Lecanorine, smooth, not flexuose, persistent or excluded, prominent or distinct, concolourous with the surrounding thallus; with a thin, inner rim visible between disk and thalline margin. Disk: Plane, brownish red, epruinose, pruina scarce. Parathecium (proper excipulum): White, not inspersed with crystals. Amphithecium (thalline excipulum): Present; crystals small, abundant, dissolving in 10% KOH; photobiont abundant. Amphithecial Cortex: Present, indistinct, eucortical, (35)-40-50-60 µm wide, basally widened, (50)-60-80-100 µm wide at the base; inspersed with oil droplets. Episamma: Without episamma. Epithecium: Apical cells slightly swollen, brownish red, insoluble in 10% KOH. Hymenium: Not inspers. Hypothecium: White; not inspers. Ascospores: c. 8 per ascus, broadly ellipsoid, (9.5)-10-12-14.5-(15) µm long, (6)-6.5-7-8.5-(9) µm wide; wall thick, distinctly differentiated into primary and secondary wall, not ornamented. Secondary Metabolites: Atranorin and hopane-6a,22-diol, of the following substance class(es): orcinol depsides and (tri-)terpenoids. Spot Tests: Upper surface: C –, PD –.