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Verrucaria fusca Pers.  
Family: Verrucariaceae
Verrucaria fusca image
Othmar Breuss  
Thallus: irregularly granular-areolate, thin, sometimes becoming subgelatinous when wetted, partly with a dark fimbriate prothallus areoles: convex to flattened, 0.1-0.2 mm thick, dispersed to crowded in scattered patches or coalescing to form an uneven irregularly areolate crust surface: brown to brown-black, dull, rough anatomy: paraplectenchymatous throughout, with a pigmented uppermost cell layer, without an epinecral layer, with mycobiont cells 5-8 µm in diam. and algal cells 6-10 µm in diam. distributed over most of the thallus Perithecia: prominent, hemispherical, inconspicuous as of the same color as thallus; exciple: brown, 0.2-0.3 mm wide, subglobose or depressed-ovate ; involucrellum: hemispherical, with a thin thalline covering, appressed to the exciple, extending down as far as the base of the perithecium, 30-60 µm thick, even in thickness or somewhat broadening or thinning toward base; periphyses 15-25 µm long, rather thickish asci: clavate, 70-80 x 16-22 µm, 8-spored ascospores: hyaline, simple, ellipsoid, 17-22 x 8-10 µm Pycnidia: unknown Spot tests: all negative Secondary metabolites: none detected. Substrate and ecology: epilithic, on calcareous rocks and caliche World distribution: Europe and North America Sonoran distribution: Arizona, southern California, and Baja California. Notes: Verrucaria fusca resembles V. memnonia but the latter has smaller spores. Verrucaria inornata is also similar but has thinner involucrella, colorless exciples, and larger spores.
Verrucaria fusca image
Verrucaria fusca image
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