Ecology: Biotroph; lichenized; episubstratic; substrate metalliferous. Thallus: Areolate (primarily areolate), continuous throughout, dispersed along the periphery, continuous in the centre, or dispersed throughout and distinctly seperate; lobes rounded at the front; separate thallus parts (.2)-.3-1-(1.4) mm thick. Thallus Size and Differentiation: Large; main branches basally not constricted or basally constricted. Thallus Outline: Margin concolorous the thallus centre. Upper Surface: Red (rust-red), convex; not sorediate; without thalloconidia thalloconidia. Medulla: Iodine reaction in Lugol's solution positive. Ascocarps: Soon sessile, slightly constricted or strongly constricted at the base, 15-100 per cm², .5-.8-(2) mm in diam.. Margin: Not flexuose, persistent or excluded, 40-80 µm wide, rarely epruinose. Disk: Plane, weakly convex, or strongly convex, black, epruinose, without an umbo. Exciple: Hyphae radiating outwards; 2.5-4 µm wide; grey; not inspersed with crystals or inspersed with crystals. Epithecium: Apical cells green or turquoise, cell pigment reacting with 10% HCl (in H2O), cell pigment HCl+ blue. Hymenium: 45-65 µm high; white or pale green; iodine reaction: Lugol’s positive. Subhymenium: 20-40 µm high; white or pale green. Hypothecium: Brownish yellow or brown. Ascospores: Ellipsoid, (7)-8.3-10.1-(14) µm long, (4.5)-5.2-6.1-(7) µm wide; wall not ornamented. Conidia: (6)-7.5-10.5-(12) µm long. Secondary Metabolites: Not detected or present, stictic acid or porphyrilic acid. Spot Tests: Medulla: K – or rarely + yellow, C –, PD – or rarely + orange; ascocarp margin (in section): K –, C – disk: C – hypothecium: K – epihymenium: K – or + green.