Ecology: Terricolous; growing sun-exposed or partially shaded. Thallus: Crustose, squamulose; lobes ascending; branches densely aggregated, scattered, loosely aggregated, or clustered; separate thallus parts 2-15 mm long; 2-4 mm wide. Primary Thallus: Evanescent or persistent. Secondary Thallus: Present, of determinate growth; stipe 8-80 mm high, 1-4 mm wide, lime green or olive, homogeneously coloured or mottled, corticate, with intact or longitudinally split wall. Thallus Outline: Margin crenulate or dissected. Upper Surface: Areolated, lime green or olive, smooth; not sorediate. Lower Surface: White. Lower Cortex: Absent. Ascocarps: Forming apical at the tip of branches or stipes or along the margin of scyphi, moderately abundant, 1-2 mm in diam.. Disk: Weakly convex to strongly convex; brown. Conidiomata: Formed on podetia or on margins of scyphi. Secondary Metabolites: Fumarprotocetraric acid and protocetraric acid, of the following substance class(es): ß-orcinol depsidones. Spot Tests: Upper surface: K (negative), C , PD + red.