Global occurrence: Eurasia – Asia Extratropical | Eurasia – Europe | Arctic. Substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts. Life habit: lichenized (mutualistic with algal photobionts). Thallus: crustose (crustaceous) – endosubstratal, inconspicuous, immersed | crustose (crustaceous) – episubstratal – unspecified; continuous, diffuse, effuse | granular, granulose, granulate; [th] upper surface: white(ish); [th marginal and upper surface] specific structures: absent; [th] morphol substructures (eg areoles, lobes, branches) upper surface: granulose, granular. Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (low) 0.5 (high) 0.8; [ascm, if apoth] disc, mazaedium: concave; [ascm, if apoth] disc, mazaedium: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | black(ish) brown; [ascm, if apoth] disc, mazaedium: epruinose; [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: yellow(ish) (if dull: buff) | brown(ish) (if pale: fawn, tan; if mid: cinnamon); [ascm] paraphyses/-oids: present; [ascm] paraphyses/-oid cells width [µm]: (median) 1.0. Asci: unitunicate; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): absent. Ascospores: (min) 4.0 (low) 6.0 (high) 8.0; [asp] shape: ellipsoidal | ovoid, ovate; [asp] length [µm]: (low) 28.0 (high) 50.0; [asp] width [µm]: (low) 15.0 (high) 25.0; [asp] septa: present, muriformly septate – muriform, submuriform; [asp] transversal septa: (low) 7.0 (high) 17.0; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: thick, distincly halonate. Secondary metabolites: absent. Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid. Secondary photobionts (eg in cephalodia): absent.