Global occurrence: Eurasia – Asia Extratropical | Eurasia – Europe | Americas – North America (incl Mexico) | Arctic. Substrate: rock – siliceous, siliciferous, acidic | rock, stones, pebbles – unspecified. Life habit: lichenized (mutualistic with algal photobionts). Thallus: crustose (crustaceous) – episubstratal – unspecified; compartimentized – areolate, squamulate; [th] upper surface: grey(ish); [th upper surface]: epruinose; [th marginal and upper surface] specific structures: absent; [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 0.3 (high) 1.0 (max) 2.5; [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane | verrucose, warted | subsquamulose; [th] medulla amyloidity: absent. Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (low) 0.5 (high) 1.0; ascoma: sessile, superficial; [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded | subconvex, slightly convex | convex; [ascm, if apoth] disc, mazaedium: black(ish); [ascm, if apoth] disc, mazaedium: epruinose; [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: black(ish); [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: blue(ish) brown (violet brown) | red(dish) brown (if pale: orange brown) | black(ish) red; [ascm] paraphyses/-oids: present; [ascm] paraphyses/-oid cells width [µm]: (median) 1.5; [ascm] paraphyses/-oid apical cells width [µm]: (low) 2.5 (high) 3.0; [ascm] epihymenium, epithecium: green(ish) grey | brown(ish) green (olivaceous, olive green). Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: amyloid with widening axial body towards the apex (= Lecanora-, Parmelia-, Rinodina-types etc). Ascospores: (median) 8.0; [asp] shape: ellipsoidal | oblong, oblong-obtuse; [asp] length [µm]: (min) 9.0 (low) 12.0 (high) 15.0 (max) 17.0; [asp] width [µm]: (low) 4.0 (high) 6.0 (max) 6.5; [asp] septa: absent – spore lumen unilocular, monolocular; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent. Conidiomata: absent | present. Conidia: filiform, vermiform; [co] length [µm]: (low) 22.0 (high) 30.0. Secondary metabolites: present, miriquidic acid. Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid. Secondary photobionts (eg in cephalodia): absent.